Gained as part of the main story at the end of Chapter 3.
Gained as part of the main story, near the start of Chapter 1.
Gained as part of the main story at the start of Chapter 4
Found in Chapter 4, and missable if you don't trigger the statue that grants it.
The game does direct you to activate the statue though, so this could be considered as part of the main story.
In addition, other statues appear later in the game - but I don't know if they count for this or not.
Gained as part of the main story at the end of Chapter 5.
Gained as part of the main story at the end of Chapter 7
Gained as part of the main story at the end of Chapter 7
Gained as part of the main story at the end of Chapter 9
Gained as part of the story, just after the end of Chapter 9.
Gained as part of the story at the end of Chapter 10.
Since it's an ending, it *technically* requires multiple playthroughs - but you are able to continue from the main menu to claim both within a few minutes of each other.
Since it's an ending, it *technically* requires multiple playthroughs - but you are able to continue from the main menu to claim both within a few minutes of each other.
Missable since it requires you to either drop the cube from a high place (which are very rare, and seem to not work on the Steam version), or crush in inside some platforms (and a valid location only appears in 1 chapter).
Missable since it can only be gained in a handful of spots (primarily the triple platform room in Chapter 6)